The Causses" is a near 3400 km2 large plateau located in the south of France. Here the first dinosaur footprints where found in 1935. After this, this area has yielded an ever-increasing number of ichnites now in excess of 500 specimens. These latter, 15 to 50 cm long, tridactyl or tetradactyl footprints of generally biped animals, were discovered at the surface of Hettangian to lower Sinemurian dolomite layers within 4 distinct stratigraphic units. The 35 sites bearing ichnites are located on the plateau margin. For the first time, morphologic characters studied through descriptive statistic methods with the usual parameters and classical Student and Snédecor tests, allowed us, to divide the whole set of biped traces into 6 ichnospecies. Their definitions are further constrained by multivariate statistical results using Principal Component Analysis (PCA), Factor Analysis of correspondances (FAC) and Discriminant Analysis (DA). All have confirmed the morphologic observations. So that now, the following taxa are identified : Grallator variabilis, G. lescurei, G. sauclierensis, G. minusculus, Eubrontes giganteus, Dilophosauripus williamsi, cf. Moraesichnium, Orníthopus fabrei nov ichnosp. The more immediately visible differences relate to the interdigital II-IV divarication and the digit length ratio. To this panel, we must add Batrachopus deweyi and shapes suggesting Trisauropodichnus and/or Anomoepus. Among all ichnite associations described in the lower Liasic, the New England assemblage presents the most affinities with ours. It shows the ichnotaxa Grallator, Dilophosauripus, Eubrontes, Batrachopus without forgetting Ornithopus fabrei nov. ichnosp. which is close to Ornithopus gallinaceus from the Massachusetts and Connecticut basins. On comparing the present early Jurassic ichnofauna of the Causses with the ones of the Middle and Upper Triassic formations of the eastem border of the Massif Central (France), it appears that tridactyl footprints become more and more numerous and large from Triassic to Early Jurassic. In the Causses, these latest are prevalent but in Quercy (France), Poland, Italy, USA, they are also associated with Omithopoda, Thyreophora and Sauropoda ichnites. Footprint areas considered here were generaly under an arid climate. Animals that passed by were heavy and bulky possible Megalosaur trackmakers, and lighter and slender Coelophysids or Ceratosaurs. For all, these areas were pathways as the orientations of the trackways seem point out. The directions followed by these reptiles were without any important variation during the Hettango-Sinemurian stages. These areas were also used from time to time by Crocodilomorpha and may be tetradactyl (I-IV) bipedal avian Theropods. However, the number of such trackways in sites, sometimes substantial, should not lead us to overestimate the trackmakers populations. These last were probably relatively moderately abondant in this inter-supratidal swamp environment. In the Causses, ichnites are connected with former algo-laminated deposits (Algal mats) which were rapidly hardened by means of calcitisation of cyanobacteria. The result has been a moderate depth of footprints; autopodia disturbing only a few cm of the carbonate substrate. Other fossils have been discovered : invertebrates with thin bivalve and gastropod shells, crustaceans tests and plants. These latter suggest the existence of paleomangroves like environments but also continental vegetation periodically overruning the swamp environment during regression/transgression cycles. At these times, wooded parts of it, could become protecting, feeding, resting and nesting places.

  

Herein we describe a small but relatively diverse assemblage of fossil fishes derived from the lower Oligocene (Rupelian) Red Bluff Clay at site AMo-9 in Monroe County, Alabama, USA. Identified amongst the remains are 15 unequivocal taxa representing 11 families within five orders, and one additional taxon represents an unknown order and family. Taxa identified include Eostegostoma sp., Otodus (Carcharocles) sp., Mitsukurinidae/Carchariidae indet., Macrorhizodus praecursor, Galeorhinus sp., Negaprion gilmorei, Physogaleus sp., “Sphyrna” sp., Galeocerdo sp., cf. “Aetobatus” sp., Sphyraena sp., Xiphiorhynchus kimblalocki, Xiphiorhynchus sp., Cylindracanthus ornatus, and C. rectus. Several additional fossils could not be identified beyond Lamniformes, Carcharhiniformes, and Teleostei, but they likely belong to one of the identified taxa within this paleofauna. All of the fishes previously reported from the Red Bluff Clay within the entirety of the Gulf Coastal Plain of the USA are otolith-based, and each of the 15 unequivocal taxa reported herein are important new records for this lithostratigraphic unit. In particular, the Eostegostoma sp. and Xiphiorhynchus spp. specimens represent the first occurrences of these taxa in Alabama. The specimens of C. ornatus, Eostegostoma sp., and X. kimblalocki are stratigraphic and temporal range extensions from the middle and late Eocene into the Rupelian Stage of the Oligocene. Other described taxa may represent transitional forms between those described from the late Eocene and late Oligocene within the region. This study provides a tantalizing preliminary view into faunal transitions that occurred amongst marine fishes across the Eocene/Oligocene boundary within the Gulf Coastal Plain of the USA. 

The present study describes in detail the anterior limb osteology of a molossid chiropteran of the genus Tadarida, from Céreste, a Stampian locality in the Apt-Forcalquier Oligocene basin already known for its fishes, plants and insects.

A comparision with the Miocene forms T. srehlini from Saint-Géraud localities and T. sp. from Württemberg, also with the recent forms T. teniotis and Eumops perotis, does not show any clear morphological differences between the Tertiary and Recent Tadarida, indicating a rather noticeable anatomical stability, not exceptionnal indeed among Chiropterans. The Céreste fossil exhibits however slightly primitive wing proportions if compared to the Saint Gérand Aquitanian species.

Several remarks deal with the peculiar relationships between the ecology of the molossids and their kind of fossilisation, frequently associated with sedimentary facies of the lacustrine type.
  

The locality of Dams (Quercy, southwestern France) has yielded two fossil assemblages, one from the late Eocene and another from the early Oligocene, making it one of the few localities with infillings across the Eocene-Oligocene transition. At least 24 taxa (13 mammals, 11 snakes) have been identified in this locality. Study of the lissamphibian remains from Dams yields an Eocene and an Oligocene assemblage, with a total of eight taxa. The Eocene assemblage includes two unnamed salamandrine species, one unnamed pelobatid species and one pyxicephalid species (Thaumastosaurus). The Oligocene assemblage includes two unnamed pleurodeline species, one salamandrine species (Salamandra sansaniensis) and an unnamed pelobatid species. Among the eight taxa from Dams, one Eocene salamandrine and one Oligocene pleurodeline are identified for the first time in the Quercy. A review of the lissamphibians from the Quercy area identifies eleven taxa for the Late Eocene (MP19) and eight taxa for Early Oligocene (MP22), with a major turnover at the Eocene-Oligocene transition. This turnover occurs in a time of major climatic changes, with a significant decrease in temperature and precipitation and concurrent increase in seasonality in Europe, likely affecting specialized taxa.
  

A new hypothesis for the phylogenetic relationships of recent anomalurids and graphiurids is proposed, based on information from evolutionary lineages of Paleogene European rodents, particularly Gliridae, and Eocene Algerian Zegdoumyidae. Differences in first occurrences, in paleogeography, and in infraorbital structure in glirids (protrogomorphy and pseudomyomorphy) and graphiurids (hystricomorphy) separate Graphiuridae from Gliridae (Graphiurinae is here raised to family rank). Similar considerations, and dental morphology, suggest that Anomaluridae (appearing in the late Eocene) and Graphiuridae (appearing in the Pliocene) are related to early Eocene Zegdoumyidae. 

Previous chronological arrangements of South American Quaternary land mammal faunas are appraised on the basis of current geological and paleontological data. Three South American late Pliocene-Pleistocene land mammal ages are conventionally recognized, from oldest to youngest, the Uquian, Ensenadan, and Lujanian ; all are defined on Argentine faunas.

     The Uquian is based fundamentally and historically on the fauna from the Uquía Formation in Jujuy Province, northwestern Argentina. Important known formations in Argentina yielding Uquian Age faunas include the sub-surface Puelche Formation (or Puelchense) near the city of Buenos Aires, and the Barranca de Los Lobos and Vorohué Formations between Mar del Plata and Miramar, Buenos Aires Province. A tentative subdivision is propos-ed for the Uquian into three subages based on knowledge of the Mar del Plata-Miramar sequence, from oldest to youngest, the Barrancalobian, Vorohuean, and Sanandresian. In Argentina the Uquian is presently marked by the first known record of Scelidodon, Hydrochoeropsis, Ctenomys, Canidae, Ursidae, Gomphotheriidae, Equidae, Tapiridae, Camelidae, Cervidae, and the last known record of Thylatheridium, Thylophorops, Dankomys, Eumysops, Pithanotomys, Eucoelophorus, Hegetotheriidae, Sparassocynidae, and Microtragulidae.

The Ensenadan Age is based on the fauna from the Ensenada Formation near the city of Ensenada, Buenos Aires Province. In Argentina the Ensenadan is marked by the first known record of Lomaphorus, Neothoracophorus, Plaxhaplous, Cavia, Lyncodon, Lutra, Galera, Smilodon, Dicotyles, Lama, Vicugna, the last known record of Orthomyctera, and the only known record of Brachynasua.

     Typícal beds of late Lujanian Age in Argentina consist of fluvial deposits occupying stream channels, and shallow basins, often incised into beds of early Lujanian (i.e. Bonaerian of early workers) and Ensenadan Age. The Lujanian Age is based on a fauna from beds along the Rio Luján, about 65 km west of the city of Buenos Aires, Buenos Aires Province. The Lujanian in Argentina is marked by the first record of Equus, Chlamyphorus, and Holochilus, and the last record of Megatherioidea, Glyptodontoidea, Arctodus (=Arctotherium), Smilodon, Litopterna, Notoungulata, Proboscidea, Equidae, Morenelaphus, and Palaeolama.

   These land mammal ages are often difficult to recognize in other South American countries. The compositions of South American Pleistocene faunas vary with the environment. Some taxa were widely distributed in fossil deposits throughout the continent, but their occurrences need not reflect synchroneity. This is a result of changing climates and habitats in time. Consequently, proposed intracontinental correlations need confirmation based on magnetostratigraphy and a radioisotope time scale. Paleontologic characterizations of these land mammal ages (i.e. first and last record, and guide fossils) are useful for much of Argentina, but extensions to most of the other parts of South America are at best tenuous.

The majority of known non-Argentine Pleistocene faunas are believed to be Lujanian in age. Possible non Argentine early Pleistocene (Uquian) faunas include Ayo Ayo and Anzaldo in Bolivia, and Cocha Verde in southern Columbia. A possible middle Pleistocene (Ensenadan or early Lujanian) fauna is the Chichense of Ecuador. Paleomagnetic and radioisotopic date (MacFadden et al., 1983) clearly indicate that the greater part of the Tarija fauna (Bolivia) is Ensenadan in age.

  The end of the Pleistocene and beginning of the Holocene in South America is marked by extinction of nearly all large mammalian herbivores and their specialized large predators. Radiocarbon age determinations suggest that large scale extinctions of megafauna occurred between 15,000 and 8,000 yrs. B.P. (years before present). 

Morphological descriptions are given of Eocene mammals from the locality Toru Ajgyr (NEKyrgyzstan) that were excavated in 1997 and 1998 in a cooperation between the Martin-Luther-University Halle (Germany), the Zoological Institute in St. Petersburg (Russia) and the Seismological Institute in Bishkek (Kyrgyzstan). The species found belong mostly to perissodactyls, as Lophialetes sp., Teleolophus sp. and brontotheres. The primitive ungulate family Olseniidae is represented by a complete foot skeleton of cf. Olsenia sp. In addition, postcranial materials of Gobiatherium mirificum (Dinocerata) and of artiodactyls have been collected and are described herein. Based on mammals, the locality is part of the Asian Land Mammal Age Arshantan and is stratigraphically equivalent with the Bridgerian Land Mammal Age in North America and with the lower and middle Geiseltalian of the European Middle Eocene. 

The locality of Fons 1, one of the fossiliferous outcrops in the late Eocene limestones of Fons-outre-Gardon (Gard), has yielded varied remains of mammals. The specimens were prepared by dilute acetic acid attack on the rock and by impregnation with an acrylic resin.
This fauna, rich with about 20 species, can be included in the biochronologic zone of Euzet. The Palaeotheridae (sensu Simpson 1945), represented by 6 species, occupy a predominant place; Plagiolophus annectens is particularly abundant, comprising 55% of all the specimens found.
The abundance of this species has permitted a detailed comparative study with the corresponding form from Euzet. The quantitative tests brought out a slight but highly sígnificant difference between the average dimensions of the teeth in the two populations. Moreover and in spite of the feeble rate of evolution in the Plagialophus dentition, diverse archaic particularities can be seen which permit the conclusion that the form from Fons 1 is a little the older. This conclusion is corroborated by observations on the other palaeotherids, particularly Lophiotherium aff. cervulum, whose dental evolutionary stage is intermediate between those of the corresponding forms known from Robiac and Euzet.
A new species of Pachynolophus is described, P. garimondi n.sp., the last representative known of the genus. lts teeth are distinguished by their size, a great heterodonty, and the weakness
of their cingula. 

Tetraceratops insignis is known from a single, crushed skull from the Lower Permian of Texas. Its unique proportions and osteological details gained central meaning in the question of the origins of Therapsida since this early synapsid has been determined as the oldest and less derived therapsid. Apart from Tetraceratops, the ‘mammal-like’ Therapsida and their sister, the pelycosaur-grade Sphenacodontidae, are separated by one of the longest ghost lineages in tetrapod fossil record. However, the minor, though well justified critique faced insistent publication regarding the therapsid hypothesis. A carefull re-evaluation of the holotypic skull reveals that therapsid traits cannot be supported, including a rejection of the formerly supposed adductor shelf in the temporal fenestra. Increased understanding of ‘pelycosaur’ character variation underlines a haptodontine-grade or, less likely, sphenacodontid position for Tetraceratops. 

Confident attribution of bipedal tridactyl dinosaur tracks to theropods or ornithopods can be challenging. Here we describe trackways produced by tetanuran dinosaurs, previously attributed to hadrosaurs, from Coahuila State, northeastern Mexico. Multiple trackways headed in the same direction suggest gregarious behaviour in these late Campanian theropods.  

Based on material found in about 15 localities the relationships of the genera Microparamys and Glirarus have been studied. One new genus, two subgenera and three species [Microparamys (Sparnacomys) chandoni n. subgen. and n. sp., Microparamys (Pantrogna) russelli n. subgen., Eoglirarus wildi n. gen. and n. sp., Gliravus meridionalis n. sp.] as well as the publication
of numerous new facts concerning species previously reported, support the phyletic scheme proposed. The latter shows that the origin of the Gliravinae is to be sought among the very small and still rather poorly known Microparamys species of the early Eocene. Gliravus: robiacensis can be considered as the common ancestor to different lineages not only of Glirarus but also of modern genera (Peridyromys, Glirudinus and Microdyromyx), at the origin of which Peridyromys micio, although difficult  to interpret, occupies a similar place.

The stratigraphic conclusions permit more detail in the chronologie succession of the localities studied. The paleoecologic and biogeographic aspects lead one to the problem of the oligocene "Grande Coupure" through the study of this group.

  

A comparative bone histological study of the three known genera of the endemic European ornithopod dinosaur family, Rhabdodontidae, is presented here in an ontogenetic context. Investigated specimens were assigned to different ontogenetic stages based exclusively on the histological indicators of osteologic maturation during diametrical bone growth; an entirely size-independent method as opposed to most previous studies. Qualitative comparison of bone histology of corresponding ontogenetic stages and elements among the three valid rhabdodontid genera, Mochlodon, Zalmoxes, and Rhabdodon, revealed some consistent patterns. Genus specific histological differences within Rhabdodontidae are most expressed between Rhabdodon and the Mochlodon-Zalmoxes clade. These indicate a prolonged phase of fast growth and a less constrained cyclicity in the growth dynamics of Rhabdodon, as opposed to the slower and more regulated growth strategy reflected in the bones of Mochlodon and Zalmoxes. These genus specific differences are consistent with the phylogenetic interrelation of the genera and are most probably related to the pronounced differences in body size. However, when compared to other ornithopods, most detected histological features in rhabdodontids do not seem to reliably reflect either phylogenetic relations or body size. A notable common feature of all rhabdodontid genera irrespective of body size is the ontogenetically early onset of cyclical growth and secondary remodelling; a pattern that more resembles the condition found in derived ornithopods than that described in more basal taxa which are closer relatives of rhabdodontids. The recognition of taxon-specific histological patterns as well as patterns indicative of ecological and thereby functional traits clearly requires more accurate, preferably quantitative evaluations.   

Some new material of percrocutas from the late Miocene of Axios valley (Macedonia, Greece) is studied. They have been found in the locality of "Pentalophos 1" (PNT). The material has been described and compared with the known late Miocene percrocutas of Eurasia. This comparison indicates that it can be identified as Dinocrocuta gigantea (SCHLOSSER, 1903). A maxilla of a percrocuta, named ”Hyaena" salonicae, was found in the same area (Andrews, 1918). "Hyaena" salonicae is smaller than the PNT material. It is also compared with other material from Eurasia while its taxonomic and age problems are discussed. It belongs to Dinocrocuta and shows close relationships with D. robusta and D. senyureki; its age can be considered as late Vallesian-early Turolian. The age of the locality PNT is also discussed and a possible Vallesian age is proposed for it. 

NA 

The recent discovery of several mandibular toothplates of a chimaeroid fish at a dig in the area of the Landover Mall, near Landover, Prince Georges County, Maryland, brings to our attention a new species of the chimaeroid, Ischyodus.
 Although superficially reminiscent of the European chimaeroid toothplates of Ischyodus thurmanni (cf. text-fig. 2), I. williamsae nov. sp. probably has a more closer relationship to I. bifurcatus CASE, 1978, of the Upper Cretaceous of New Jersey and Delaware. 

Les Pistes de Vertébrés du Stormberg Supérieur ("Trias terminal à Rhétien"), ou Quthingien

Si les zones du Stormberg inférieur se sont révélées contenir de nombreuses traces, surtout dans les faciès dits "Molteno moyen et supérieur", représentant apparemment la base du Keuper, il est frappant de voir pratiquement l'ensemble de cette grosse faune "Molteno" disparaître avec la fin de cette période, que nous avons appelée le "Maphutsengien".

Dès les premières zones du Stormberg supérieur, que nous nommons le"Quthingien" la zoocénose et la phytocénose, en même temps que les données d'ensemble manifestées par l'environnement, sont modifiées. Nous ne verrons plus guère de dépôts marécageux à flore riche et variée, parfois même luxuriante. Les fougères elles-mêmes ont disparu. Elles sont remplacées par de maigres plantes, aux feuilles très souvent filiformes qui paraissent témoigner d'un climat continental. Le sol est devenu de plus en plus rouge, avec des variations latérales beaucoup plus accusées. Les fleuves amenant des galets des monts du "Grand Sud" ont tari. La faune va en subir les conséquences. Certaines des espèces se révèleront sautillantes ou coureuses, pour un grand nombre plus légères et pour la quasi-totalité d'apparence carnivore ou entomophages, les phytophages devant se contenter d'un régime ingrat,difficile ou à tout le moins irrégulier,les dépôts le montrent.

C'est dans ces conditions que s'inaugure notre Etage nouveau,quelque peu discordant sur les zones A/5, A/6 ou A/7 du Stormberg inférieur (Maphutsengien). Le Stormberg supérieur (ou Quthingien) commence avec le paléopaysage remarquable dit de Moyeni, que nous allons maintenant étudier, typologiquement, avec ses homologues du même âge. Quelques 38 types d'animaux tous nouveaux vont défiler à nos yeux lors de la zone de base de cet Etage, ou zone B/1.

L'on nous avait proposé d'intituler ce Ile Tome de la série : "La grande Dalle de Moyeni et ses homologues. Paléo-spectacles, scènes et paysages animaux au Lesotho à l'approche du Trias finissant". Nous avons préféré garder le sous-titre plus haut, peut-être plus prosaïque.

Un llle Tome est en préparation : "Les développements ultérieurs et terminaux de la faune du Gondwana".

  

Redécouvertes en 1968 dans le bois du Bretou, les deux poches à phosphorites qui ont livré cette faune sont parmi les plus méridionales du plateau quercynois. De faible profondeur, huit mètres, les cavités sont de petite taille et font partie d'un complexe de sept fissures réparties dans les bois du Bretou. Les autres trous n'ont livré aucun fossile, soit parce qu'ils avaient été intégralement vidés de leur sédiment, soit que des venues d'eau en interdisent l'accès.
Dans les deux poches fossilifères, proches de quelques mètres, le sédiment que l'on rencontre en placage sur les parois présente un aspect particulier. Il s'agit en effet d'une brèche très indurée, où la calcite domine, et contenant assez peu de sédiments sidérolithiques. Les ossements contenus dans cette brèche sont assez nombreux, très souvent brisés. Les dents isolées y sont relativement rares. Exploitées séparément, les deux poches ont livré des faunes identiques, aucune différence chronologique ne pouvant être décelée entre elles. Les formes rencontrées dans une seule des cavités sont rares mais la poche la plus riche est celle dénommée Le Bretou 2.
L'intérêt de la faune récoltée réside dans son ancienneté. On sait en effet que H.G. Stehlin avait signalé dans les collections du Quercy des restes attribuables à Lophiodon lautricense, mais la provenance exacte de ces fossiles reste inconnue. Au Bretou si nous n'avons pas rencontré à ce jour le Lophiodon, il n'en demeure pas moins que l'âge de la faune récoltée doit être proche de celui de la faune de Robiac. 

A cheek tooth recently unearthed in the Lopar Sandstone unit, of late Eocene age, in the northern part of Rab Island, Croatia, is one of the very few Eocene mammalian remains found in the Adriatic area. Thorough comparison of this tooth with those of Old-World Palaeogene mammalian orders suggests that it is a M3 belonging to an embrithopod afrothere. The specimen is referred to as Palaeoamasia sp. This genus was formerly known only in Eocene deposits of Anatolia but with close relatives in Romania among Palaeoamasiinae. The geographical distribution of this subfamily perfectly matches the recently-named Balkanatolian landmass, which experienced in-situ evolution of endemic mammals prior to the Grande Coupure event that occurred around the Eocene–Oligocene transition. This last event is characterised by massive Asian immigration in Western Europe and the supposed extinction of many endemic Central and Western European mammals, including Palaeoamasiinae.
  

Lalieudorhynchus gandi gen. nov. and sp. nov. is a new caseid synapsid from the Permian of the Lodève Basin, Occitanie, France. This new taxon is represented by a partial but well-preserved postcranial skeleton, and is characterized by the following apomorphies: a transverse section of the sacral and anterior caudal neural spines with a very thin keel-like process anteriorly, a slender dorsal tip of the dorsal and caudal spines, a narrow distal end of the first sacral rib, a fossa on triceps process of metacoracoid, and a very large distal tarsal 1 of same width than the astragalus, with nearly all sides being shallowly concave.
The skeleton corresponds to a sub-adult individual that was excavated from the La Lieude Formation dated as Roadian-Capitanian (Guadalupian). A sedimentological and taphonomical analysis of the type locality, together with preliminary osteohistological observations, suggest that this new French caseid was rather aquatic, as already hypothesised for other large forms.
A phylogenetic analysis of caseids is performed to test the position of this new taxon and to better understand the evolution of the clade: interestingly, Lalieudorhynchus gandi gen. nov. et sp. nov. is closer to the NorthAmerican “Cotylorhynchus” hancocki than to the other French caseids Ruthenosaurus and Euromycter from the Artinskian of the geographically closer Rodez Basin. These two last caseids document the Artinskian radiation of the clade, which remained diverse until Olson’s extinction. Caseids survived, as Lalieudorhynchus is one of the youngest representatives of the clade, and may have used novel ecological strategies to access their vegetarian food sources.

  

A new artiodactyl, Diacodexis antunesi n.sp., is described from the early Eocene of Silveirinha, Portugal. Comparisons are made with Diacodexis gazini GODINOT, 1978, D. varleti SUDRE et al., 1983, D. cf. varleti from Paris Basin sites, D. sp. from Dormaal and from localities in Spain and England, D. secans from North America and D. pakistanensis from Asia; affinities and evolutive tendencies are discussed. The presence of Diacodexis in the locality of Silveirinha confirms the very early Eocene age of the latter. As Diacodexis antunesi appears ta be more primitive than D. gazini from Rians (early Eocene of France), it lends corroboration to the interpretation (essentially based previously on condylarths) of the Silveirinha assemblage as the oldest Eocene fauna known in Europe and supports the hypothesis that early artiodactyls migrated from Europe to North America.